Must Sexually Selected Traits Be "Honest Indicators" Under the Runaway

Must sexually selected traits be "honest indicators" under the runaway sexual selection model?

A) Yes, traits under sexual selection always must be costly.
B) No, female preference alone is sufficient under runaway selection.
C) Yes, otherwise genetic linkage could not be established.
D) No, the model only requires female preference and male traits are coded on the same chromosome.
E) B and C

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