In the Evolution of Whelk-Eating Behaviour in Crows, Which of the Following

In the evolution of whelk-eating behaviour in crows, which of the following was optimized through natural selection?

A) the average number of drops required to break the shell
B) the average height a bird flew to drop a shell
C) the average total energy used to break shells
D) the average size of the shells dropped by the birds
E) the average thickness of the shells dropped by the birds

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