What Would Happen If Mitochondria Were Treated with a Proton

What would happen if mitochondria were treated with a proton gradient uncoupler,such as 2,4-dinitrophenol?

A) Electron transfer would stop.
B) Complex I would become reduced, and complexes III and IV would become oxidized.
C) Protons would be pumped by the mitochondrial electron transport chain, although no ATP would be synthesized.
D) Reducing equivalents, in the form of NADH, would no longer be consumed.

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